Fall 2022

Developmental Biology

Listed in: Biology, as BIOL-221

Moodle site: Course


Katerina Ragkousi (Sections 01 and L01)


How can a single cell, the fertilized egg, give rise to all the specialized cells of an adult? What gives rise to biological form? What is the molecular logic of the pathways that progressively refine cellular identities? How do cells "talk" to one another so as to coordinate their behaviors as embryos develop form and function? How can parts of an organism be regenerated with only the appropriate regions remade, structured identically to the missing ones? How does a stem cell differ from a non-stem cell? How can genetically identical organisms be cloned? This course will offer an integrative study of the development of animals, leading to the formulation of the principles of development, including an introduction to experimental embryology and developmental physiology, anatomy, genetics and "evo-devo." Laboratory work explores embryonic development and regeneration in amphibians, sea urchins, nematodes, flatworms, fruit flies, fish, and chickens. Three classroom hours plus three hours of laboratory per week.Requisite: BIOL 191. Not open to first-year students. Limited to 16 students. Fall 2022. Professor Ragkousi.

How to handle overenrollment: First preference to senior majors, then seniors in related disciplines, then junior majors, etc.

Students who enroll in this course will likely encounter and be expected to engage in the following intellectual skills, modes of learning, and assessment: an emphasis on group work, critical reading and discussion of primary literature, in-class exams, independent research, lab notebook keeping and lab visit in and out defined practical hours for observation of embryonic development.


Lab Science Course


2022-23: Offered in Fall 2022
Other years: Offered in Fall 2014, Spring 2016, Fall 2022, Spring 2024